Organismal Biology #21
SIMPLE ANIMALS: SPONGES AND CNIDARIA

Kingdom Animalia includes all organisms that develop from a hollow ball of cells called a blastula. Most animals have well-developed motility.
The simplest animals include the sponges (Porifera) and the Cnidaria.
Sponges are unsymmetrical or radially symmetrical, with many cell types but no distinct tissues; their bodies contain numerous pores and sharp protective spicules.
Coelenterates (phylum Cnidaria) are radially symmetrical, with two tissue layers (ectoderm and endoderm) surrounding an all-purpose gastrovascular cavity.

Phylum Porifera (sponges): Aquatic animals with radial symmetry or irregular shapes. Water enters by many incurrent pores that often lead to a central cavity. Water may exit by an excurrent opening (osculum). No distinct tissues, but many cell types:
  • Epidermal cells (pinacocytes): Outside lining
  • Porocytes: Barrel-shaped pore cells
  • Choanocytes: Flagellated "collar cells" that keep water flowing
  • Amoebocytes: Several kinds of amoeboid cells, including some that secrete sharp spicules.
  • Spicules: sharp needles or more complex shapes embedded within sponges, functioning in support and as a defense against predators. May be composed of silica, calcite, or horny protein.
Tissue-level organization:   A tissue is a group of similar cells and their products, built together (structurally integrated) and working together (functionally integrated).
You may think of a sponge as built of a single tissue. Types of sponges include:
  • Calcispongia (or Calcarea):   small sponges with needle-like spicules made of calcite (CaCO3 ). Symmetry usually radial.
  • Hyalospongia (or Hexactinellida):   deep-water "glassy" sponges (often very beautiful) with spicules (usually six-pointed) made of silica (SiO2 ). Symmetry usually radial.
  • Demospongia:  Horny or "true" sponges (about 90% of all known species) with spicules made mainly of horny protein ("spongin," similar to keratin) that often surrounds a needle-like core of silica. Irregular shapes (symmetry lost).  Traditional "bath sponges" belong here.
Sponges and Cnidaria

Phylum Cnidaria (coelenterates): Aquatic animals with two tissue layers (outer ectoderm and inner endoderm) separated by a jelly-like mesoglea; and an all-purpose gastrovascular cavity with a single opening (mouth). Tentacles surround mouth and have stinging cells (cnidocytes) containing stingers (nematocysts). Two major body forms:
Polyp: mouth directed upward, mesoglea thin, animal usually attached;
Medusa: free-swimming "jellyfish" with thick mesoglea; mouth directed downward.
  • Class Hydrozoa: Life cycle includes both asexual polyps and sexually reproducing medusae (usually small). Solitary or colonial; some colonies have many types of individuals interconnected.
  • Class Scyphozoa: Solitary "jellyfish" with dominant medusa stage.
  • Class Anthozoa: The largest class, including sea anemones and corals. Polyp stage dominant; no medusa. Mouth extends inward to form a tubular pharynx. Solitary or colonial.
Phylum Ctenophora ("comb jellies"): A small group of marine animals with biradial symmetry (like a two-armed pinwheel), 2 large tentacles, and 8 comb-like rows of cilia.



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