Organismal Biology #14
ORIGIN OF EUCARYOTES
Performance Objectives:
Eucaryotic cells have nuclei surrounded by double-layered membranes. The several chromosomes each contain proteins as well as DNA. The cytoplasm of eucaryotic cells contains many organelles not found in procaryotic cells. Much evidence indicates that eucaryotic cells originated by symbiosis. In particular, mitochondria and plastids were once separate organisms.

Procaryotic vs Eucaryotic cells:
Procaryotic vs Eucaryotic cell types

Eucaryotic cells are cells with true nuclei, each containing a nucleolus and surrounded by a nuclear envelope.
  • Chromosomes are usually separate, multiple, and linear; each contains protein as well as DNA.
  • Cytoplasm contains contractile proteins (actin, myosin), making possible cytoplasmic streaming (cyclosis), amoeboid locomotion using pseudopods, and ingestion of food by phagocytosis.
  • Many types of cytoplasmic organelles are present, including membrane organelles (like endoplasmic reticulum) and "9 + 2" arrangements of microtubules. Mitochondria (and plastids in plant cells) contain their own DNA.

Theory of endosymbiosis: Most biologists now believe that eucaryotic cells originated when small, energy-producing procaryotic cells lived inside larger cells and became mitochondria by intracellular symbiosis (endosymbiosis). Plastids may have arisen the same way. Similar origins for other cell parts have been proposed but are less widely accepted. Perhaps host cells originally phagocytized the energy-producing mitochondria; then natural selection favored those host cells that maintained the mitochondria as an energy source instead of digesting them.
  • Evidence for endosymbiosis comes from the fact that both mitochondria and plastids have two membranes: the outer membrane resembles the membranes of the eucaryotic host cell, but the inner one resembles procaryotic cell membranes instead.
  • Mitochondria and plastids possess their own DNA and are self-replicating.
  • Bacteria are known whose metabolic abilities are similar to those of the postulated host cell, while other bacteria have the enzymes of the Krebs cycle and are thus comparable in ability to mitochondria.
  • REVIEW:         Study guide and vocabulary

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